Gentians

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Gentians
Clusius gentian (Gentiana clusii)

Clusius gentian ( Gentiana clusii )

Systematics
Asterids
Euasterids I
Order : Enzianartige (Gentianales)
Family : Gentian Family (Gentianaceae)
Tribe : Gentianeae
Genre : Gentians
Scientific name
Gentiana
L.

The gentians ( Gentiana ) are a genus of flowering plants in the family of the Gentian family (Gentianaceae). The 300 to 400 species, which occur almost worldwide, mainly thrive in the mountains of the temperate zones of the northern hemisphere , but also in the Andes . Some types are used to make schnapps and to obtain medicinal products. Few species and varieties are used as ornamental plants .

description

Illustration of the Eastern Alpine gentian ( Gentiana pannonica ) from Germany's flora in images based on nature
Long-life bracts, fruit and seeds of the Alpine gentian ( Gentiana alpina )
Seeds of Gentiana triflora var. Japonica

Vegetative characteristics

Gentian species grow as annual or perennial herbaceous plants . The ascending to independently erect stems are grooved or angular. In some perennial species there are generative and vegetative shoot axes .

The opposite or rarely whorled leaves are in basal rosettes or distributed on the stem. The simple leaf blades have a smooth leaf margin.

Generative characteristics

The flowers are laterally or terminally, individually or usually several in zymotic inflorescences or sometimes in small groups or whorls.

The hermaphroditic flowers are radial symmetry and usually five, rarely four or six to eight-fold with a double flower envelope . The green sepals are fused to about half of their length. The calyx teeth are thread- to egg-shaped and have a raised median nerve. The usually five, rarely four or six to eight petals are often long-tubular, inverted-cone-shaped, bell-shaped, plate-shaped, jar-shaped or rarely wheel-shaped (probably only in Gentiana lutea ). Usually the corolla tubes are much longer than the corolla lobes. There are plicae between the corolla lobes. The petals can have very different shades of blue or are yellow to whitish. There is only a circle with four or five, rarely four or six to eight stamens ; they are fused with the corolla tube. The stamens do not protrude beyond the petals. The stamens are more or less winged at their base. The anthers are usually free, they are rarely connected. There are five to ten glands at the base of the ovary. The sedentary or located at a long Gynophor ovary is upper constant and usually at least single chamber in the upper region. The stylus is usually short and linear, or more rarely long and thread-shaped or absent. The scar lobes are free or fused, bent back, usually elongated to linear, rarely spread out and rounded.

The capsule fruits are cylindrical to ellipsoid and winged, or more or less narrowly obovate (narrow ellipsoid in Gentiana winchuanensis ), have two fruit valves and contain many seeds. The seeds can be winged or wingless. The seed coat is tiny, reticulate, wrinkled or areolate (= field).

ecology

In most species of gentian, the flowers close when the sky is cloudy and the air is cool, even when subjected to vibrations from hail, rain and strong wind.

Systematics and distribution

Herbarium evidence : holotype of Gentiana pumila subsp. delphinensis

The genus Gentiana was in 1753 by Carl von Linné in Species Plantarum , 1, pp. 227-232. set up. Gentiana lutea L. was used as a lectotype in 1913 by Nathaniel Lord Britton and Addison Brown in An Illustrated Flora of the Northern United States, Canada, & the British Possessions. 2nd Edition. Volume 3, p. 8. Synonyms for Gentiana L. are: Calathiana Delarbre , Chondrophylla A.Nelson , Ciminalis Adans. , Dasystephana Adans. , Ericala Gray orth. Var., Eurythalia D.Don orth. Var., Favargera Á.Löve & D.Löve , Gentianodes Á.Löve & D.Löve , Holubogentia Á.Löve & D.Löve , Kuepferella M.Laínz , Kurramiana Omer & Qaiser , Mehraea Á.Löve & D.Löve , Pneumonanthe Gled. , Qaisera Omer , Tretorhiza Adans. , Ulostoma G.Don .

The botanical genus name Gentiana is the Greek name for species of this genus and has been shown to have been used since 50 to 100 AD. According to Dioscoroides and Pliny the Elder, it is derived from the Illyrian king Genthios (180–168 BC ), who is said to have been the first to recognize the healing properties of gentian species (probably Gentiana lutea ).

Koch's gentian
( Gentiana acaulis )
Clusius gentian
( Gentiana clusii )

The genus Gentiana belongs to the subtribe Gentianinae from the tribe Gentianeae within the Gentianaceae family . Earlier, the genres were gentianella ( Gentianella ) gentianopsis ( Gentianopsis ) and hair throat ( Comastoma ) to a large genus gentian ( Gentiana counted sl).

The genus Gentiana is divided into ten or eleven sections:

  • Section Gentiana sect. Asclepiadeae (Grossh. Ex Gagnidze) Tzvelev (for example: Gentiana asclepiadea )
  • Section Gentiana sect. Calathianae (for example: Gentiana bavarica )
  • Section Gentiana sect. Chondrophyllae Bunge (for example: Gentiana arisanensis )
  • Section Gentiana sect. Ciminalis (for example: Gentiana acaulis )
  • Section Gentiana sect. Cruciata Gaudin (for example: Gentiana cruciata )
  • Section Gentiana sect. Dolichocarpa T.N.Ho (example: Gentiana sedifolia )
  • Section Gentiana sect. Frigidae Kusn. (for example: Gentiana algida )
  • Section Gentiana sect. Gentiana (for example: Gentiana burseri )
  • Section Gentiana sect. Isomeria Kusn. (for example: Gentiana depressa )
  • Section Gentiana sect. Kudoa (Masamune) Satake & Toyokuni ex Toyokuni (for example: Gentiana arethusae )
  • Section Gentiana sect. Pneumonanthe Gaudin (for example: Gentiana affinis )

In Europe, around 35 species of Gentiana thrive, mainly in the Alps . The following species occur in Central Europe : Koch's gentian or silicate bell-shaped gentian ( Gentiana acaulis ), Alpine gentian ( Gentiana alpina ), narrow-leaved gentian ( Gentiana angustifolia ), swallow-wort gentian ( Gentiana asclepiadea ), Bavarian gentian ( Gentiana bavarica ), Short-leaf gentian ( Gentiana brachyphylla ), Villars-gentian ( Gentiana burseri subsp. Villarsii ), Clusius-gentian or lime-bell-gentian or stemless gentian ( Gentiana clusii ), cross-gentian ( Gentiana cruciata ), Tauern-gentian ( Gentiana frigida ), Karawanken Enzian ( Gentiana froelichii ) with the subspecies: Silvia-Zenari gentian ( Gentiana froelichii . subsp zenarii ) Ligurian gentian ( Gentiana ligustica ), yellow gentian ( Gentiana lutea ), alpine gentian ( Gentiana nivalis ), Round-leaved gentian ( Gentiana orbicularis ), Eastern Alpine Gentian or Pannonian Gentian or Hungarian Gentian ( Gentiana pannonica ), Lung Gentian ( Gentiana pneumonanthe ), Lying Gentian ( Gentiana prostrata ), Low he gentian or cute gentian ( Gentiana pumila ), spotted gentian ( Gentiana punctata ), purple gentian ( Gentiana purpurea ), Rostan's gentian ( Gentiana rostanii ), Triglav gentian ( Gentiana terglouensis subsp. terglouensis ), Schleicher's gentian ( Gentiana terglouensis subsp. schleicheri ), hose gentian ( Gentiana utriculosa ), spring gentian or “Schusternagerl” ( Gentiana verna ). The well-known trivial name "stemless gentian" describes both the Clusius gentian and the Koch gentian .

The genus Gentiana comprises 300 to 400 species worldwide, which predominantly thrive in the mountains of the temperate zones of the northern hemisphere , but also in the Andes , Gentiana species are absent in Africa . There are about 13 species in Mexico.

There are 300 to 400 species in the genus Gentiana worldwide
Narrow-leaved gentian ( Gentiana angustifolia )
Swallowwort gentian
( Gentiana asclepiadea )
Short-leaf gentian ( Gentiana brachyphylla )
Gentiana clausa inflorescence
Cross gentian ( gentiana cruciata )
Narrow-leaved gentian ( Gentiana decumbens )
Dinaric gentian ( Gentiana dinarica )
Tauern gentian ( Gentiana frigida )
Karawanken gentian ( Gentiana froelichii )
Yellow gentian ( Gentiana lutea )
The hybrid Macauleys Enzian (
Gentiana × macauleyi ) created by horticultural crossing of Gentiana farreri and Gentiana sino-ornata
Flowering in detail of Gentiana newberryi
Snow gentian ( gentiana nivalis )
Gentiana occidentalis subsp. corbariensis
Lung gentian
( Gentiana pneumonanthe )
Spotted Gentian ( Gentiana punctata )
Gentiana scabrida var. Scabrida
Gentiana septemfida var. Lagodechiana
Gentiana triflora var. Japonica
Gentiana triflora var. Montana
Hose gentian ( Gentiana utriculosa )
Spring gentian ( Gentiana verna )

symbolism

Austrian 1 cent coin

Many types of gentians have the pure blue flower color, which is rare in the plant kingdom, which is why these gentians have become a symbol of loyalty.

The gentian is depicted on the Austrian euro cent coins , as well as on the Romanian one-leu note and the Swiss five-franc coin .

The German Federal Railroad used the designation “ Blauer Enzian ” for a long - distance express train and later Trans-Europ-Express between Hamburg and Munich , the route of which was later extended to Austria.

use

The "rhizomes" of some gentian species have been used since the Middle Ages to make schnapps , as medicines and to stimulate appetite. In contrast to the blue flowering gentians depicted on many bottle labels, the large species ( tall herbaceous gentians) , especially the yellow gentian ( Gentiana lutea ), are used for distilling schnapps , in addition, purple gentian ( Gentiana purpurea ), spotted gentian ( Gentiana) are used punctata ) and the Hungarian gentian ( Gentiana pannonica ).

Gentians are also used as a bitter substance ( amarum ). The most bitter "natural" substance in the world, amarogentin , is extracted from the roots of certain species . Amarogentin is still clearly noticeable even in a dilution of one to 58 million.

With many types of gentian, the above-ground parts also taste bitter and are therefore avoided by grazing cattle. This is one reason why species of gentians have been able to keep well on the grazed alpine meadows.

All European gentian species are under nature protection , which means that they may neither be picked nor dug up. For this reason, attempts are currently being made to cultivate the gentian required for making schnapps, and a large part of the drugs required for making schnapps and medicines is already being cultivated and harvested as a special crop in fields.

Varieties of some species and hybrids are used as ornamental plants .

Misleading trivial names

The Indian yellow gentian (Kuru, Katuka), a bitter medicinal herb of the West Himalayas ( Picrorhiza kurroa Royle ex Benth .; Plantaginaceae ) and the closely related plant 胡黃蓮 hú huáng used in traditional Chinese medicine (TCM) do not belong to the genus lián , Picrorhiza scrophulariiflora Pennell (Plantaginaceae).

swell

Individual evidence

  1. a b c d e f g h i j k l m n o p q r s t u v w x y z aa ab ac ad ae af ag ah ai aj ak al am an ao ap aq ar as at au av aw ax ay az ba bb bc bd be bf bg bh bi bj bk bl bm bn bo bp bq br bs bt bu bv bw bx by bz ca cb cc cd ce cf cg ch ci cj ck cl cm cn co cp cq cr cs ct cu cv cw cx cy cz da db dc dd de df dg dh di dj dk dl dm dn do dp dq dr ds dt du dv dw dx dy dz ea eb ec ed ee ef eg eh ei ej ek el em en eo ep eq er es et eu ev ew ex ey ez fa fb fc fd fe ff fg fh fi fj fk fl fm fn fo fp fq fr fs ft fu fv fw fx fy fz ga gb gc gd ge gf gg gh gi gj gk gl gm gn go gp gq gr gs gt gu Ting-nung Ho, James S. Pringle: Gentianaceae. Gentiana Linnaeus. P. 15–93 - the same text online as the printed work , In: Wu Zheng-yi, Peter H. Raven (Ed.): Flora of China. Volume 16 - Gentianaceae through Boraginaceae , Science Press and Missouri Botanical Garden Press, Beijing and St. Louis, 1995, ISBN 0-915279-33-9 .
  2. a b c d data sheet at the Gentian Research Network .
  3. ^ A b Eckehart J. Jäger, Friedrich Ebel, Peter Hanelt, Gerd Müller: Rothmaler - Excursion flora of Germany: Herbaceous ornamental and useful plants . Springer, 2016, ISBN 978-3-662-50420-8 , pp. 406-410 ( Gentiana on pp. 406-410. In the Google book search).
  4. Linné 1753 first publication scanned on biodiversitylibrary.org .
  5. ^ A b Gentiana at Tropicos.org. Missouri Botanical Garden, St. Louis
  6. a b c d e f g h i j k l m n o p q r s t u v w x y z aa ab ac ad ae af ag ah Gentiana in the Germplasm Resources Information Network (GRIN), USDA , ARS , National Genetic Resources Program. National Germplasm Resources Laboratory, Beltsville, Maryland.
  7. David Aeschimann, Konrad Lauber, Daniel Martin Moser, Jean-Paul Theurillat: Flora alpina. An atlas of all 4500 vascular plants in the Alps . tape 2 . Haupt Verlag, Bern, Stuttgart, Vienna 2004, ISBN 3-258-06600-0 , p. 10-24 .
  8. ^ Manfred A. Fischer, Wolfgang Adler, Karl Oswald: Excursion flora for Austria, Liechtenstein and South Tyrol . 2nd, improved and enlarged edition. State of Upper Austria, Biology Center of the Upper Austrian State Museums, Linz 2005, ISBN 3-85474-140-5 .
  9. a b c d e f g h i j k l m n José A. Villarreal – Quintanilla, A. Eduardo Estrada – Castillón, Diana Jasso de Rodríguez: El género Gentiana (Gentianaceae) en México - The genus Gentiana (Gentianaceae) in Mexico. In: Polibotánica. Volume 27, México, 2009. online.
  10. a b c d e f g h i j k Gentiana in the Red List of Threatened Species of IUCN 2018-1. Retrieved on 2018-11-08.
  11. a b c d e f g h i j k l m Chih-Hsiung Chen, Jenn-Che Wang: Gentianaceae. In the Digital Flora of Taiwan.
  12. a b c d e f g Chih-Hsiung Chen, Jenn-Che Wang: Revision of Taiwanese Gentiana - Revision of the genus Gentiana L. (Gentianaceae) in Taiwan. In: Botanical Bulletin of Academia Sinica. Volume 40, 1999, pp. 9-38.
  13. a b c d e f g h i j k Karol Marhold: Gentianaceae. In: Euro + Med Plantbase - the information resource for Euro-Mediterranean plant diversity . 2011, accessed November 4, 2018 .
  14. Eitel-Friedrich Scholz: The gentian does not only bloom blue. undated , oeav-obergailtal.at .
  15. Identified taste sensors for the world's bitterest natural substance. (No longer available online.) German Institute for Nutritional Research , formerly the original ; Retrieved March 20, 2013 (press release).  ( Page no longer available , search in web archivesInfo: The link was automatically marked as defective. Please check the link according to the instructions and then remove this notice. @1@ 2Template: Dead Link / www.dife.de  
  16. Picrorhiza kurroa. Spektrumverlag: Lexicon of medicinal plants and drugs . Picrorhiza kurroa Royle ex Benth (Plantoginaceae). In: Ammon scrap: Picrorhiza kurroa Royle ex Benth (Plantoginaceae). In: Medicinal plants of Ayurvedic and Western medicine . 2012, p.
     418 ( limited preview in Google Book search). .
    Picrorhiza Kurroa (Hu Huang Lian) chineseherbshealing.com

Supplementary literature

  • Xaver Finkenzeller, Jürke Grau: Alpine flowers. Recognize and determine (=  Steinbach's natural guide ). Mosaik, Munich 2002, ISBN 3-576-11482-3 .
  • Hansjörg Küster: History of the landscape in Central Europe. Munich 1999, ISBN 3-406-45357-0 .
  • T.-N. Ho, S.-W. Liu: A worldwide monograph of Gentiana. Science Press 2001.

Web links

Commons : Enziane ( Gentiana )  - album with pictures, videos and audio files