violet

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violet
Grove violets (Viola riviniana)

Grove violets ( Viola riviniana )

Systematics
Rosids
Eurosiden I
Order : Malpighiales (Malpighiales)
Family : Violet family (Violaceae)
Subfamily : Violoideae
Genre : violet
Scientific name
viola
L.

Violets or violas ( Viola ) are a genus of plants in the violet family (Violaceae). Most of the approximately 500 species thrive in the temperate zones of the earth. The centers of biodiversity are in North America , the Andes, and Japan . However, they can also be found in Australia and Tasmania. Well-known species are pansies , horned violets and fragrant violets .

description

Illustration of viola gracilis
Illustration of the grove violet ( Viola riviniana )

Appearance and leaves

Violet species grow as annual or biennial or mostly perennial herbaceous plants , as well as, rarely, subshrubs with very changing habitus. Often rhizomes are formed as permanent organs. More or less long, creeping to upright, above-ground shoot axes can be formed or absent.

The leaves can all stand together at the base or be arranged alternately distributed on the stem axis. The leaves are divided into a petiole and a leaf blade. The leaf blades are simple or divided. The leaf margin is smooth or serrated. The remaining, small to large, sometimes foliage -like stipules are free or more or less long fused with the petioles and often ciliate or fringed and then often have brown glands at their ends.

Some viola species have leaves that are +/- dense, mostly hairy on the underside and ciliate on the edge and base. The hairs consist of simple, single-celled or single-row trichomes , which in some species are thickened towards the end. So far, only glandular villi are known of external glands. They sit on the ends of the stipules, on the teeth of the leaves and on the tips of the sepals, and consist of a rather short, thick stem and a multicellular, secreting head. The glandular villi of the stipules form mucus at an early stage and cover the developing leaf with their secretion. The mucus emerges only at individual points of the cuticle and can occur repeatedly because the subcuticular mucus formation continues. A regeneration of the cuticle of these mucous glands does not take place, but only the formation of very resistant, so-called border membranes. Excretion of lime has been observed on the marginal teeth of some species (e.g. Viola scandens ).

blossoms

The flowers stand individually in the leaf axils on flower stalks that have two bracts . Often the flowers are dimorphic, the cleistogamic ( remaining closed) flowers develop later than the chasmogamous (opening).

Opened three-lobed fruit capsule of the field pansy ( Viola arvensis )
Seeds of Viola lactea (left, dark) and Viola odorata (right, light)

The hermaphrodite flowers are zygomorphic and five-fold with a double flower envelope . The almost identical five free or almost free sepals usually have herbaceous ears (appendages) at their base. The five free petals are clearly unequal. The lowest petal is the largest and spurred at its base. There is a circle with five stamens . The free stamens are relatively short. The upright anthers are free or usually form an envelope around the ovary. The two lower anthers protrude into the spur and have spur- or wart-like, nectar-secreting appendages at their base. Three carpels have become a top permanent, single-chamber ovary grown. Each ovary contains many anatropic ovules in parietal placentation. The almost upright or mostly more or less curved downward stylus is more or less thickened or sometimes gradually narrowed upwards and is smooth or has different appendages. The scar is shaped differently depending on the species.

The color of the petals is responsible for the color of the flowers, which in the family are mostly yellow to red, purple and blue, more rarely whitish, brownish or +/- black. The often intense yellow hues are formed by small pigment granules that occur in various amounts in the colorless cell sap of the epidermal cells . The granules consist e.g. B. in Viola primulaefolia from quercetin , a plant pigment from the flavone group related to tannins . In Viola tricolor and Viola arvensis , however, they consist of rutin (= Violarutin or Violaquercitrin), a glycosidic dye of the formula C-27 H-30 O-sixteenth The blue color tones are caused by anthocyanin dissolved in the cell sap of the epidermal cells , the color of which is red, purple or blue depending on the acid, neutral or alkaline reaction of the cell sap. A wide variety of blue hues can often be found in immediately adjacent cells. If there are cells without colored cell sap in the epidermis between the blue cells, light blue nuances arise. In the pansy varieties, the black colors of individual parts of the petals are based on a combination of deep blue anthocyanin solution and yellow-red pigment granules. The deep blue- violet varieties contain the dye violanin .

Fruits and seeds

The loculicidal capsule fruits open with three elastic flaps that are keeled or thickened on the underside.

The spherical to egg-shaped seeds have a light to dark, mostly smooth surface. There may be an aril . The seeds contain abundant endosperm and a straight embryo with two very thick, plano-convex cotyledons ( cotyledons ).

Locations

The great majority of the species belong to the northern temperate zone, in the hotter regions of the world they inhabit the mountains; numerous species are characteristic endemisms of the South American Andes; few are found in the Brazilian mountains , in tropical Africa and in the Cape Province ; nine species are endemic to the Hawaiian Islands; eight species occur only in Australia (including Tasmania and New Zealand), of which some belong, including Viola hederacea Labill. , the Antarctic flora; one species, Viola papuana W.Becker & Pulle , is endemic to New Guinea.

As a rule, the violets inhabit areas with an annual rainfall of over 600 mm.

Due to the extraordinary variety of forms and widespread use, Wilhelm Becker had postulated a great age for the genus. In Europe, there are two sections among the 15 sections: Melanium and Nominium . Becker knew 75 European violets in 1925.

The altitude distribution of the genus ranges in Yunnan up to 4030 meters, Morocco 3600 meters, East Africa 1200 to 3350 meters, Madagascar 1200 to 2700 meters, Malacca 900 to 1200 meters, Colorado up to 4050 meters.

Distribution, evolution and systematics

Flower of Viola adunca
Eastern Alps pansy ( Viola alpina )
Field pansy ( Viola arvensis )
Yellow violet ( Viola biflora )
Long-tailed violet ( Viola calcarata )
Viola calcarata subsp. zoysii
Mont Cenis violets ( Viola cenisia )
Teide violets ( Viola cheiranthifolia )
Hill violet ( Viola collina )
Viola crassa subsp. borealis
Tall violet ( Viola elatior )
Violet calamine pansy ( Viola guestphalica )
Flowers of Viola lactea
Sudeten pansy ( Viola lutea ssp. Sudetica )
Wonder Violet ( Viola mirabilis )
Scented violets ( Viola odorata )
Marsh violet ( Viola palustris )
Viola pedata with palmate leaves
Flowers of Viola philippica
Feather violets ( Viola pinnata )
Low violet ( Viola pumila )
Forest violets ( Violareichenbachiana )
Grove violets ( Viola riviniana )
Viola stipularis at La Soufrière in Guadeloupe
Wild pansy ( Viola tricolor )
Valdieri violet flowers ( Viola valderia )
Habit, leaves and flowers of Viola vallicola
Habit, leaves and flowers of Viola violacea
Habit, leaves and flowers of Viola walteri
Habit, leaves and flower of Viola yezoensis
Cultivar of the garden pansy ( Viola × wittrockiana )

distribution

The genus Viola is practically coherent in the northern hemisphere . Exceptions are the Central Asian steppes, the Tibetan highlands and the East Asian deserts. In the southern hemisphere only mountain regions are populated: in South America the arc of the Andes to Tierra del Fuego and the Falkland Islands (exclave in the Brazilian mountainous region); in Africa the Ethiopian Highlands, the East African Mountains, Madagascar and the Cape Province; all of South and Southeast Asia (the Ghats in India and the mountains of Sri Lanka) and in the Australian region the high mountains of New Guinea, the mountains of East Australia, Tasmania and New Zealand.

The Flora Europaea lists 92 species, the flora of the Soviet Union 106 species, and the flora of Japan 48 species. In the New World there are 47 species in the northeastern USA and 25 species in the western USA.

The global center of diversity is the Mediterranean region (especially south-east Europe with around 60 species, the Apennine peninsula with around 40 species) and the subtropical areas of South America with 55 species. Other centers of diversity are the islands of Japan (approx. 48 species), northern East Asia and Ussuria (35 species).

As a microthermal clan, the genus Viola has been able to colonize extensive habitats with various locations in the Holarctic area. The great abundance of forms and wide radiation of the species were promoted by the special reproductive biology (periodic alternation between foreign and self-pollination).

evolution

The species-rich viola genus is cosmopolitan . The evolution to today's species was promoted in particular by polyploidy . Polyploidy ranges from diploid (2x) to at least octadecaploid (18x) in the lines of the genus, with both very young (neo-) and older (meso-) polyploidy being observed. A total of 620 species are recognized, of which 580 species have been subdivided into 16 morphologically, chromosomically and geographically defined groups in preliminary sections. Only three of the sixteen sections have chromosomes whose base number is consistent with diploidy . According to knowledge gained from biogeography , karyology and phylogeny , a South American origin is suggested, from where the genus has spread to the northern hemisphere and beyond. Based on fossil seed discoveries, settlement in Eurasia is dated 17-18 million years ago.

Taxonomy

The first publication of Viola was made in 1753 by Carl Linnaeus in Species Plantarum , 2, pp 933-937. Type species is Viola odorata L. Synonyms for Viola L. are Erpetion Sweet and Mnemion Spach .

External system

The genus Viola belongs to the tribe Violeae in the subfamily Violoideae within the family Violaceae .

Internal system

The genus Viola is divided into several sub-genera, for example Chamaemelanium , Dischidium , Erpetion , Melanium .

Species and their distribution

The genus Viola contains over 500 to 550 species :

Hybrids :

The following are no longer included in the viola genre :

use

Some types and varieties of violets are used as ornamental plants . Some species and varieties serve as medicinal and kitchen plants and are used for cosmetic purposes.

Violets as ornamental plants

In parks and gardens in beds or planters , the most common ornamental plants are the scented violet , the horned violet and, probably the most commonly cultivated, the garden pansy , as well as the grove violet with the shape 'Purpurea' (often incorrectly referred to as viola labradorica ), the Peony violets with the varieties 'Albiflora' and 'Freckles' as well as some alpine species for the rock garden. The Parma violet is rarely found in gardens these days.

On the other hand, the Alpine violets and the African violets known as indoor plants do not belong to the genus of violets ( Viola ) .

Violets as medicinal and culinary plants

Scented violets, especially March and Parma violets, have long played an important role in herbalism. Except in the field of naturopathy , especially aromatherapy , they are no longer used. Fragranced and horned violet flowers, like pansy flowers, can also be used in culinary terms and are, for example, one of the ingredients of the original Pear Helene recipe. The well-known violet pastilles are flavored with synthetic violet fragrances ( ionones ). Liqueur flavored with violets is called crème de violette and has been made at least since the early 19th century.

Violets in perfumery

The flowers of March and Parma violets have been used in cosmetic products for centuries. The natural violet blossom absolute has practically disappeared today and is being replaced by the synthetic ionones by the cosmetics industry . Violet leaf absolute (from Viola odorata ), on the other hand, continues to be widely used in perfume production. The so-called "natural violet perfume" has also been extracted for a long time from the so-called violet root / iris root, which consists of dried rhizomes of various types of iris .

etymology

The scientific name Viola is an independent borrowing of the Latin viola (e.g. from Virgil, Columella, Pliny) and is probably the diminutive of the ancient Greek íon or víon , which already occurs in Pindar and Homer and in antiquity with the mythological Io in Has been linked. An exact equivalent of the Latin word can also be found in the Greek name Iólē , or older Viólā , the mythological daughter of King Eurytus of Oichalia on Euboia.

The German name Veilchen can be traced back directly to the Latin name viola , which became fiol in Old High German .

The name pansy is said to be based on a comparison: the two upper petals, usually without a drawing, are the stepdaughters, the conspicuously drawn petals on the side are the natural daughters and the large lower petals with the large sap mark are the mother.

Violets in mythology, customs and literature: see scented violets .

Cultural history

Various mythological stories about the violet were already known to the Greeks, who associated the violet with Io. Ioplokos or ioplokamos were called women with dark curls and iostephanos meant violet- wreathed , which in Pindar's name stands for the city of Athens. Legends explain the name in such a way that Ionic nymphs offered this flower to Jon when Athens was founded, or that Zeus offered it to his lover, the king's daughter Io, as a sweet dish. The ancient authors distinguish three groups of violets: the blue (black or purple), the actual ion ( Viola odorata ), also ion to melan (black violet with Theophrastus), ion porhyrun (purple violet with Dioscurides) and viola purpurea (with the Romans) so called; the white violet, ion to leukon or leukoion, with the Romans viola alba (Pliny), pallens viola (Virgil), leucoium (Columella); and the yellow violet, leukoion melinon (Dioscurides), viola lutea (Pliny), viola crocea (Albertus Magnus).

swell

  • Hans Melchior: The phylogenetic development of the Violaceae and the natural relationships of their genera. In: Repertorium specierum novarum regni vegetabilis - Feddes Repertorium , Botanische Beihefte. 36, 83-125, Berlin 1925.
  • Yousheng Chen, Qiner Yang, Hideaki Ohba, Vladimir V. Nikitin: Violaceae. Viola - online. , In: Wu Zheng-yi, Peter H. Raven, Deyuan Hong (Eds.): Flora of China . tape 13 : Clusiaceae through Araliaceae . Science Press / Missouri Botanical Garden Press, Beijing / St. Louis 2007, ISBN 978-1-930723-59-7 , pp. 138 (English). . (Sections Description and Systematics)
  • David Henriques Valentine , Hermann Merxmüller , Alexander Schmidt: Viola L. In: TG Tutin, VH Heywood, NA Burges, DM Moore, DH Valentine, SM Walters, DA Webb (eds.): Flora Europaea . Volume 2: Rosaceae to Umbelliferae . Cambridge University Press, Cambridge 1968, ISBN 0-521-06662-X , pp. 270–282 (English, limited preview in Google Book Search).

Individual evidence

  1. Hans Melchior: Violaceae. In: A. Engler, K. Prantl 1925: The natural plant families - 21 . Jena 1925. pp. 331-332.
  2. a b Hans Melchior: Violaceae. In: A. Engler, K. Prantl 1925: The natural plant families - 21 . Jena 1925. p. 334.
  3. Hans Melchior: Violaceae. Pp. 329-363. In: A. Engler, K. Prantl 1925: The natural plant families 21, Leipzig 1925. P. 335.
  4. Wilhelm Becker: Viola L. P. 363-376. In: A. Engler, K. Prantl 1925: The natural plant families 21, Leipzig 1925. P. 363.
  5. J. Christopher Havran, Kenneth J. Sytsma, Harvey E. Ballard Jr. (2009): Evolutionary relationships, interisland biogeography, and molecular evolution in the Hawaiian violets (Viola: Violaceae). American Journal of Botany 96 (11): 2087-2099.
  6. Hermann Meusel ; Eckehart Jäger , Stephan Rauschert, Erich Weinert 1978: Comparative Chorology of Central European Flora - Text, Volume II . Gustav Fischer, Jena. P. 292
  7. Hermann Meusel; Eckehart Jäger, Stephan Rauschert, Erich Weinert 1978: Comparative Chorology of Central European Flora - Maps, Volume II . Gustav Fischer, Jena. P. 289
  8. Thomas Marcussen, Lise Heier, Anne K. Brysting, Bengt Oxelman, Kjetill S. Jakobsen: From Gene Trees to a Dated Allopolyploid Network: Insights from the Angiosperm Genus Viola (Violaceae). In: Systematic Biology , October 3, 2014. Full text online.
  9. a b Thomas Marcussen et al. 2014: p. 85
  10. a b c d e f g h i j k l m n o p q r s t u v w x y z aa ab ac ad ae af ag ah ai aj ak al am an ao ap aq ar as at au av aw ax ay az ba bb bc bd be bf bg bh bi bj bk bl bm bn bo bp bq br bs bt bu bv bw bx by bz ca cb cc cd ce cf cg ch ci cj ck cl cm cn co cp cq cr cs ct cu cv cw cx cy cz da db dc dd de df dg dh di dj dk dl dm dn do dp dq dr ds dt du dv dw dx dy dz ea eb ec ed ee ef eg eh ei ej ek el em en eo ep eq er es et eu ev ew ex ey ez fa fb fc fd fe ff fg fh fi fj fk fl fm fn fo fp fq Viola in the Germplasm Resources Information Network (GRIN), USDA , ARS , National Genetic Resources Program. National Germplasm Resources Laboratory, Beltsville, Maryland.
  11. a b c d e f g h i j k l m n o p q r s t Yousheng Chen, Qiner Yang, Hideaki Ohba, Vladimir V. Nikitin: Violaceae. : Viola , p. 74 - the same text online as the printed work , In: Wu Zheng-yi, Peter H. Raven, Deyuan Hong (Eds.): Flora of China , Volume 13 - Clusiaceae through Araliaceae , Science Press and Missouri Botanical Garden Press , Beijing and St. Louis 2007, ISBN 978-1-930723-59-7 .
  12. a b c Matthias Erben: Cytotaxonomic investigations on south-east European viola species of the Melanium section. In: Communications from the Botanical State Collection, Munich. Vol. 21, No. 2, 1985, pp. 339-740, online .
  13. Thomas Raus: Additions and amendments to the flora of the Karpathos island group (Dodekanesos, Greece). In: Botanika Chronika. Volume 12, 1996, pp. 21-53.
  14. ^ Peter Hadland Davis (ed.): Flora of Turkey and the East Aegean Islands. Vol. 1 (Pteridophyta to Polygalaceae) . Edinburgh University Press, Edinburgh 1965, ISBN 0-85224-159-3 , pp. 531 (reprinted 1997).
  15. Richard Willfort: Health through medicinal herbs: recognition, effect u. Application. 11th edition, Trauner, Linz 1971. ISBN 3-85320-001-X .
  16. Dieter Martinetz, Roland Hartwig: Pocket book of fragrances: a lexicon from AZ. Harri Deutsch, Thun 1998, ISBN 3-8171-1539-3 .
  17. ^ H. Gams, W. Becker, 1925: Violaceae. In: G. Hegi: Illustrated flora of Central Europe. 1st ed. Volume V (1), Carl Hanser, Munich, pp. 585-668. P. 587.
  18. Helmut Genaust: Etymological dictionary of botanical plant names. 3rd, completely revised and expanded edition. Nikol, Hamburg 2005, ISBN 3-937872-16-7 (reprint from 1996).
  19. ^ H. Gams, W. Becker, 1925: Violaceae. In: G. Hegi: Illustrated flora of Central Europe. 1st ed. Volume V (1), Carl Hanser, Munich, pp. 586-587.

Supplementary literature

Identification key

Web links

Commons : Veilchen ( Viola )  - Album with pictures, videos and audio files
Wiktionary: violets  - explanations of meanings, word origins, synonyms, translations