Rock flowers

from Wikipedia, the free encyclopedia
Rock flowers
Evergreen rockflower (Draba aizoides)

Evergreen rockflower ( Draba aizoides )

Systematics
Rosids
Eurosiden II
Order : Cruciferous (Brassicales)
Family : Cruciferous vegetables (Brassicaceae)
Tribe : Arabideae
Genre : Rock flowers
Scientific name
Draba
L.

The rock florets ( Draba ), few are called hunger flowers, form a genus of plants in the cruciferous family (Brassicaceae). The 350 to 402 species are widespread in Eurasia , northwestern Africa, from North America to Mexico and across the Andes to Patagonia ; most species occur in the northern hemisphere. Most species thrive in the subarctic to arctic and / or subalpine to alpine flora .

description

Illustration of A (left) alpine rock flowers ( Draba alpina ) and B (right) snow rock flowers ( Draba nivalis )
Illustration of A (left) spring hunger flowers ( Draba verna ) and B (right) Norwegian rock flowers ( Draba norvegica )
Illustration from atlas of the alpine flora of the evergreen rockflower ( Draba aizoides )
Illustration of the ice rock flower ( Draba dubia ) from Atlas of the Alpine flora
Illustration from The fur seals and fur-seal islands of the North Pacific Ocean by Draba grandis

Appearance, indument and foliage leaves

Draba species grow as rarely annual, biennial or perennial or usually perennial herbaceous plants , sometimes the stems lignify at their base and rarely are subshrubs . The mostly upright to ascending, rarely lying to prostrate stem axes are unbranched or, mostly in the upper area, branched.

The plant parts are hairless to very different: the trichomes can be stalked, they are simple or, for example, forked, cross-shaped, star-shaped and often several of these trichome forms are present in one species.

The leaves are mostly in basal rosettes and arranged alternately on the stem, sometimes the stem is leafless. The basal leaves are usually stalked, rarely sessile and their simple leaf blades have a smooth or serrated, rarely lobed leaf margin. If stem leaves are present, then they are stalked or sessile and their simple leaf blades with a mostly wedge-shaped or rarely hollowed blade base have a smooth or toothed leaf margin.

Inflorescences and flowers

Draba species usually have umbrella-like inflorescences , the inflorescence axes of which can lengthen until the fruit is ripe and thus develop into racemose inflorescences. Sometimes bracts present. In the inflorescences there are a few to many flowers.

The hermaphrodite flowers are fourfold. The four rarely durable, upright to ascending or rarely spreading sepals are ovate or oblong, rarely elliptical. The edges of the sepals are mostly membranous. The four mostly yellow, white, seldom pink to purple or orange, seldom red petals are obovate, spatulate, lanceolate or linear, seldom circular or oblong, they are indistinctly to clearly nailed and the upper end is blunt, rounded , notched or rarely deeply divided into two parts. The petals are usually longer, rarely shorter than the sepals. Of the six stamens , four are longer and two are shorter. The stamens can be widened at their base. The anthers are ovate or oblong with a blunt upper end. The one, two or four (as the middle ones may be missing) nectar glands stand alone or are confluent. The split by a complete, wrong septum ovary contains 4 to 70 (often up to 88) ovules . The durable, cylindrical and with a length of 0.5 to 17 millimeters barely to clearly recognizable stylus ends in a cephalic scar that is simple or somewhat bilobed.

Fruits and seeds

The thin fruit stalks are upright to ascending or spreading. The bare pods or mostly silicles are ovate, lanceolate, elliptical, oblong, linear, lanceolate or nearly circular, flat or sometimes spirally rotated. The bald or fluffy hairy, paper-like fruit flaps have distinct or inconspicuous central and lateral nerves. The placental frame (replum) is rounded. The fully formed septum is membranous and translucent. In the fruit, the seeds are arranged in two rows. When ripe, the fruits open and release the seeds.

The elongated, egg or circular seeds are mostly flattened, in Draba aleutica and Draba verna only slightly flattened, and mostly wingless, but winged in Draba asterophora , Draba brachycarpa , Draba carnosula , Draba pterosperma . The tiny net-like seed coat (testa) is not slimy when wet. The cotyledons ( cotyledons ) are bent back.

Sets of chromosomes

The basic chromosome numbers are x = 6 to 12, most often 8. The degree of ploidy within the genus is very different from diploid to octadecaploid. The species examined are most common with 23% aneuploid and the second most common with a good 20% tetraploid.

Locations and exposure

Draba species thrive predominantly in alpine or boreal areas, they rarely grow in temperate and low-lying areas of the northern hemisphere. Many species of rock florets inhabit open, stony soils : rocks, crevices, rock debris or gaps in dry grass .

About 14 Draba species are listed on the IUCN's Red List of Endangered Species . Not all states register their species on the IUCN list and there are many more Draba species in the red lists of their home states. For example, nine other species are endangered in Japan.

Systematics and distribution

Taxonomy

The genus Draba was set up in 1753 by Carl von Linné in Species Plantarum , Volume 2, pp. 642-643. The generic name Draba is derived from the Greek word drabe and means "hot", "bitter" or "burning". As Lektotypusart was Draba incana L. by Mary Letitia Green in the Bulletin of Miscellaneous Information , established in 1925, p. 51 Otto Eugen Schulz published a monograph on the Draba genus in 1927 : Cruciferae-Draba et Erophila. In: Heinrich Gustav Adolf Engler : The Plant Kingdom: ... , Booklet 89, IV, p. 105.

Synonyms for Draba L. are: Aizodraba Fourr. , Coelonema Maxim. , Dolichostylis Turcz. non Cassini , Drabella Nábelek non (DC.) Fourr. , Drabopsis C. Koch , Erophila DC. , Holargidium Turczaninow ex Ledebour , Leptonema Hook. non A.Juss. nec Rabenh. nec J.Sm. , Nesodraba Greene , Odontocyclus Turcz. , Pseudobraya Korsh. , Schivereckia Andrz. ex DC. , Stenonema Hook. , Thylacodraba (Nábelek) OESchulz .

Species and their distribution

Draba is the most species-rich and taxonomically most difficult genus of the Brassicaceae family. Over 950 taxa , almost a quarter of them as infraspecific taxa, have been published. Some authors used to associate some species with the hunger flowers ( Erophila ). But the spring hunger flower ( Draba verna L. ), for example , is considered accepted in the Flora of North America 2010 and in Ingrid Jordon-Thaden 2010, since most authors, such as Al-Shehbaz, have all Erophila species in the genus Draba can be placed.

The 350 to 402 Draba species are mainly found in the northern hemisphere in Eurasia and North America as well as in North Africa. They especially thrive in arctic, subarctic , alpine and subalpine regions. Only a few species thrive in the lower altitudes of the temperate areas of the northern hemisphere. About 70 species occur in South America in the Andes from Colombia to Patagonia . There are about 121 species in North America. There are about 48 species in China , 16 of them only there.

External system

The genus Draba belongs to the tribe Arabideae within the family Brassicaceae .

In the 21st century, due to molecular genetic data, some changes were necessary in the tribe Arabideae, this also affects species that were classified in the genus Draba and some genera poor in species were reactivated or newly established:

swell

  • Ihsan A. Al-Shehbaz, Michael D. Windham, Reidar Elven: Draba , pp. 269–346 - online with the same text as the printed work , In: Flora of North America Editorial Committee (ed.): Flora of North America North of Mexico , Volume 7 - Magnoliophyta: Salicaceae to Brassicaceae , Oxford University Press, New York and Oxford 2010, ISBN 978-0-19-531822-7 . (Sections Description, Distribution and Systematics)
  • SG Aiken, MJ Dallwitz, LL Consaul, CL McJannet, LJ Gillespie, RL Boles, GW Argus, JM Gillett, PJ Scott, R. Elven, MC LeBlanc, AK Brysting, H. Solstad: Flora of the Canadian Arctic Archipelago: Draba L . online at DELTA. (Section description)

Individual evidence

  1. a b c d e f g h i j k l m n o p q r s t u v w x y z aa ab ac ad ae af ag ah ai aj ak al am an ao ap aq ar as at au av aw ax ay az ba bb bc bd be bf bg bh bi bj bk bl bm Tai-yien Cheo, Lianli Lu, Guang Yang, Ihsan A. Al-Shehbaz, Vladimir Dorofeev: Brassicaceae : Draba , pp. 66–84 - same text online as Printed work , In: Wu Zheng-Yi, Peter H. Raven, Deyuan Hong (editors): Flora of China , Volume 8 - Brassicaceae through Saxifragaceae , Science Press and Missouri Botanical Garden Press, Beijing and St. Louis, 2001, ISBN 0 -915279-93-2 .
  2. a b c d e f g h i j k l m n o p q r s t u v w x y z aa ab ac ad ae af ag ah ai aj ak al am an ao ap aq ar as at au av aw ax ay az ba bb bc bd be bf bg bh bi bj bk bl bm bn bo bp bq br bs bt bu bv bw bx by bz ca cb cc cd ce cf cg ch ci cj ck cl cm cn co cp cq cr cs ct cu cv cw cx cy cz da db dc dd de df dg dh di dj dk dl dm dn do dp dq dr ds dt du dv dw dx dy dz ea eb ec ed ee ef eg eh ei ej ek el em en eo ep eq er es et eu ev ew ex ey ez fa fb fc fd fe ff fg fh fi fj fk fl fm fn fo fp fq fr fs ft fu fv fw fx fy fz ga gb gc gd ge gf gg gh gi gj gk gl gm gn go gp gq gr gs gt gu gv gw gx gy gz ha hb hc hd he hf hg hh hi hj hk hl hm hn ho hp hq hr hs ht hu hv hw hx hy hz ia ib ic id ie if ig ih ii ij ik il im in io ip iq ir is it iu iv iw ix iy iz ja jb jc jd je jf jg jh ji jj jk jl jm jn jo jp jq jr js jt ju jv jw jx jy jz ka kb kc kd ke kf kg kh ki kj kk kl km kn ko kp kq kr ks kt ku kv kw kx ky kz la lb lc ld le lf lg lh li lj lk ll lm ln lo lp lq lr ls lt lu lv lw lx ly lz ma mb mc m d me mf mg mh mi mj mk ml mm mn mo mp mq mr ms mt mu mv mw mx my mz na nb nc nd ne nf ng nh ni nj nk nl nm nn no np nq nr ns nt nu nv nw nx ny nz oa ob oc od oe of og oh oi oj ok ol om on oo op oq or os Database: BrassiBase, Version 1.2, March 2017 of the University of Heidelberg.
  3. a b c d e f g h i j k l m n o p q r s t u v w x y z aa ab ac ad ae af ag ah ai aj ak al am an ao ap aq ar as at au av aw ax ay az ba bb bc bd be bf bg bh bi bj bk bl bm bn bo bp bq br bs bt bu bv bw bx by bz ca cb cc cd ce cf cg ch ci cj ck cl cm cn co cp cq cr cs ct cu cv cw cx cy cz da db dc dd de df dg dh di dj dk dl dm dn do dp dq dr ds dt du dv dw dx dy dz Ihsan A. Al-Shehbaz, Michael D. Windham, Reidar Elven: Draba , S. 269–346 - the same text online as the printed work , In: Flora of North America Editorial Committee (Ed.): Flora of North America North of Mexico , Volume 7 - Magnoliophyta: Salicaceae to Brassicaceae , Oxford University Press, New York and Oxford 2010, ISBN 978-0-19-531822-7
  4. ^ Ingrid Jordon-Thaden, Ihsan A. Al-Shehbaz, Marcus A. Koch: Species richness of the globally distributed, arctic-alpine genus Draba L. (Brassicaceae). In: Alpine Botany , Volume 123, Issue 2, October 2013, pp. 97-106. doi : 10.1007 / s00035-013-0120-9
  5. a b c d e f g h i j k l m n o p q r s t u v w x y z aa ab ac ad ae af ag ah ai aj ak al am an ao ap aq ar as at au av aw ax ay az ba bb bc bd be bf bg bh bi bj bk bl bm bn bo bp Draba at Tropicos.org. In: Brassicaceae . Missouri Botanical Garden, St. Louis
  6. a b c d Ingrid Jordon-Thaden: Species and Genetic Diversity of Draba: Phylogeny and Phylogeography. Dissertation at the University of Heidelberg 2010 ( full text PDF. )
  7. a b c d e f g h i j k l m n o Draba in the IUCN Red List of Threatened Species 2017. Accessed August 8, 2017.
  8. Red List of Endangered Plant Species in Japan.
  9. Enter Draba in the search mask.
  10. First publication scanned at biodiversitylibrary.org .
  11. ^ Draba at Tropicos.org. Missouri Botanical Garden, St. Louis, Retrieved August 4, 2017.
  12. a b c d e f g h i j k l m n o p q r s t u v w x y z aa ab ac ad ae af ag ah ai aj ak al am an ao ap aq ar as at au av aw ax ay az ba bb bc bd be bf bg bh bi bj bk bl bm bn bo bp bq br bs bt bu bv bw bx Entry on The Euro + Med Plantbase Project last viewed on August 8, 2017
  13. a b c d e f g h Draba in the Germplasm Resources Information Network (GRIN), USDA , ARS , National Genetic Resources Program. National Germplasm Resources Laboratory, Beltsville, Maryland., Last edited 2010.
  14. a b c d Jaakko Jalas, Juha Suominen: Atlas florae europaeae. Volume 11 Cruciferae (Ricotia to Raphanus). Pp. 80-111, Helsinki 1996, ISBN 951-9108-11-4 .
  15. a b c d e f g h i j k l m n o p Draba at Tropicos.org. In: Catalog of the Vascular Plants of Ecuador . Missouri Botanical Garden, St. Louis
  16. a b c d e f g h i Draba at Tropicos.org. In: Bolivia Checklist . Missouri Botanical Garden, St. Louis
  17. a b c d e f g h i Draba at Tropicos.org. In: Peru Checklist . Missouri Botanical Garden, St. Louis
  18. a b c Ihsan A. Al-Shehbaz, DA German, R. Karl, Ingrid Jordon-Thaden, Marcus A. Koch: Nomenclatural adjustments in the tribe Arabideae (Brassicaceae). In: Plant Diversity and Evolution , Volume 129, Issue 1, 2011, pp. 71-76. ISSN  1869-6155 doi : 10.1127 / 1869-6155 / 2011 / 0129-0044 Full-text PDF from ResearchGate .
  19. a b Ihsan A. Al-Shehbaz: A generic and tribal synopsis of Brassicaceae (Cruciferae). In: Taxon Volume 61, Issue 5, 2012, pp. 931–954. JSTOR 41679341 ISSN  0040-0262

further reading

  • Xaver Finkenzeller, Jürke Grau: Alpine flowers. Recognize and determine (=  Steinbach's natural guide ). Mosaik, Munich 2002, ISBN 3-576-11482-3 .
  • Wolfgang Adler, Karl Oswald, Raimund Fischer: Excursion flora of Austria . Ed .: Manfred A. Fischer. Eugen Ulmer, Stuttgart / Vienna 1994, ISBN 3-8001-3461-6 .
  • Ingrid Jordon-Thaden, Irina Hase, Ihsan A. Al-Shehbaz, Marcus A. Koch: Molecular phylogeny and systematics of the genus Draba sl (Brassicaceae) and identification of its closest related genera. In: Molecular Phylogenetics and Evolution , Volume 55, Issue 2, 2010, pp. 524-540. doi : 10.1016 / j.ympev.2010.02.012 online.

Web links

Commons : Rock Flowers ( Draba )  - Collection of images, videos and audio files